Education: 
B.S. Michigan State University 1959 
M.S. Michigan State University 1961 
D.V.M. Michigan State University 1961 

Academic Honors: 
Graduate of Honors College, Michigan State University 
B.S. with High Honor, Michigan State University 
Member Phi Kappa Phi, Sigma Xi, Phi Zeta 
Ciba Fellowship, 1959-1961 

Professional Experience: 
Professor Pathology, Michigan State University, 1977-present 
Associate Professor Veterinary Pathology, Washington State University, 1970-1977 
Assistant Professor Veterinary Pathology, washington State University, 1965-1970 
Guest Investigator, The Rockefeller University, 1965 
Postdoctoral Fellow, NIH, Washington State University, 1961-1965 

Research Activities: 
Studies of Genetic Disorders of Canine en Feline 
The Location and Development of Animal Models of Human Inherited Diseases 
Comparative Medicine 

Publications:  
81 publications so far 

Chapters in books and monographs: 14 so far 

Abstracts and other publications: 31 so far 

Research Grants: Total $ 3,405,648.00 

Training Grants: Total $ 1,435,567.00 

Construction Grants: Total $ 11,800,000.00 

Presentation to Canine Breed Clubs and Conferences: total 22, from 1984-1986